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PALAEONTOLOGY:
- FOSSIL THYLACINES -
(page 4)
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    Thylacinus megiriani:

    This species lived during the Late Miocene, and was discovered at Alcoota (South Quarry, Ongeva Local Fauna), in the Northern Territory.

cranial fragment of Thylacinus megiriani
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A section of the cranium of Thylacinus megiriani, including several maxillary teeth.  Courtesy: Museum of Central Australia.
    The remains of Thylacinus are very rare at Alcoota, although there were forests and a permanent water supply there during T. megiriani's time some 8 million years ago.  Palaeontologists have found many fossil bones at this site which are densely packed together from animals that died within a space of just a few years.  It is theorized that a period of very unpredictable climate occurred at Alcoota during the Late Miocene, and that there was little or no rain for several years in the area.

    Thylacinus potens:

        Thylacinus potens, whose name means "powerful thylacine", is the largest thylacinid species known to have existed.  Preceding T. cynocephalus by about 4-6 million years, its remains are known only from a single Late Miocene locality (Alcoota) near Alice Springs, Northern Territory.  This thylacine had a considerably more massive build than the modern species, and a somewhat shorter, broader skull.  Its teeth also show some slight differences in morphology, and appear to be not quite as specialized for shearing as those of T. cynocephalus (Archer 1983).

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life reconstruction of Thylacinus potens - Anne Musser
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A life reconstruction of Thylacinus potens.  The pattern of the stripes is purely speculative - as is the case with other fossil species known only from their bones, we can only theorize about their coat patterns based upon living species.  Courtesy: Anne Musser.
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    Woodburne (1967) states that T. potens is quite similar to thylacines of Quaternary (our current geological time period) age, but in its possession of "...small palatal fenestrae, the presence of a stylar cusp anterior to the metastyle of M3, the more symmetrical arrangement of the parts of the upper molars and the strong cleft in the labial outline of the upper molars...", these character states in the  Miocene species "...could be interpreted as primitive features relative to a remote dasyurid ancestry".
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    Material:  Woodburne (1967) has given a comprehensive description of the holotype specimen, a poorly preserved rostrum including a palate and damaged dentition.  The fossil was found at Paine Quarry, Alcoota Homestead, in the Northern Territory.  Some dentary fragments also exist (University of California, Berkeley, paleontological collections), but these are far less complete than the rostrum.  A few additional mandibular fragments and a small section of the premolar region of a maxilla of a second skull were found on a subsequent visit to Alcoota in 1975 (the Ray E. Lemley Expedition to Alcoota).  These new specimens (Queensland Museum) however, add little to the concept of the species as known by Woodburne (1967).

    Age:  The Alcoota Local Fauna, from the Waite Formation, is interpreted to be of Late Miocene age (approx. 7 million years old).  This fauna is dated on the basis of comparison with other dated faunas.  On this basis, it is older than the Early Pliocene local faunas, such as the Chinchilla, Bluff Downs, Bow and Hamilton faunas but younger than the array of Middle Miocene local faunas such as the Tarkarooloo, Pinpa, Ngapakaldi and Ericmas faunas (dated at around 15 million years old).  As of yet however, there is no means of establishing an absolute date for the Alcoota Local Fauna.

rostrum of Thylacinus potens (holotype)
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A palatal view of the holotype specimen of Thylacinus potens Woodburne. (Archer 1982).
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life reconstruction of Thylacinus potens - Peter Schouten
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A life reconstruction of a hunting Thylacinus potens pair.  Courtesy: Peter Schouten.
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Acknowledgement: This subsection of the Thylacine Museum has been referenced (in part) from: ARCHER, M., 1982. A review of Miocene thylacinids (Thylacinidae, Marsupialia), the phylogenetic position of the Thylacinidae and the problem of apriorisms in character analysis. In "Carnivorous Marsupials - Vol. 2" (Ed. M. Archer). Roy. Zool. Soc. N.S.W.: Sydney. pp. 445-76.
References
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