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PALAEONTOLOGY:
- FOSSIL THYLACINES -
(page 3)
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    Nimbacinus richi:

    The only other species of Nimbacinus yet described, N. richi is from the Middle Miocene deposits of Bullock Creek ("Top Site", Bullock Creek Local Fauna), Northern Territory.  The holotype specimen is a well preserved right dentary containing teeth P1 through M4.  N. richi is distinguished from N. dicksoni on the basis of some minor details of dental morphology, and its status as a distinct species is currently under debate (Wroe and Musser 2001).

mandibular molars of Nimbacinus archibaldi (holotype)
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A lateral view of molars M1-4 in the holotype dentary (lower jaw) of N. richi.  (Murray and Megirian 2000).

    Thylacinus cynocephalus:

    Fossilized remains of the modern thylacine species T. cynocephalus, dating from the Pleistocene epoch, are known from many sites across Australia, including Tasmania and Papua New Guinea.  The exceptional example shown in the photographs below is a cranium found in the Yarrangobilly Caves of Mount Kosciuszko National Park, NSW in 1969.  Notable localities for T. cynocephalus fossils include the Naracoorte Caves (SA) and Wellington Caves (NSW).  For further details about the ancient distribution of the species, see the subsection Prehistoric Range of the Thylacine.

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Pleistocene-age thylacine skull - Yarrangobilly Caves, Mount Kosciuszko National Park, NSW
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Pleistocene-age thylacine skull - Yarrangobilly Caves, Mount Kosciuszko National Park, NSW
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A Pleistocene-age thylacine skull (large male) in dorsal, palatal and lateral view.
The roof of the skull appears to have been gnawed by a rodent, possibly prior to fossilization.
Janus Cave (Yarrangobilly Caves), Mount Kosciuszko National Park, NSW.  Specimen ANWC M15266.
Courtesy: Australian National Wildlife Collection (Canberra).
Photos: International Thylacine Specimen Database Fifth Revision, 2013.
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    Thylacinus macknessi:
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    T. macknessi lived during the Early Miocene.  Its fossils were found in north-western Queensland at the Riversleigh World Heritage area (Neville's Garden Site, Neville's Garden Local Fauna).  The oldest known member of the genus Thylacinus, its species name honors Brian Mackness, a ong-time supporter of Australian vertebrate palaeontology.  The holotype specimen of T. macknessi
is a nearly complete right dentary (lower jaw) in which all teeth are present except for the incisors.  When the species was first described, only the posterior section of the jaw (bearing the last three molars) was known.  Luckily, the anterior half of the holotype specimen was found two years later in a block of limestone from the same fossil site (Muirhead and Gillespie 1995).  In some respects, this species is more specialized than more recent members of its genus, which suggests that a common ancestor of all species of Thylacinus must have been older than T. macknessi. .
Riversleigh thylacine / modern thylacine jaw comparison
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A section of the lower jaw from one of the larger species of fossil thylacines found at Riversleigh is shown here above the jaw of a modern thylacine (T. cynocephalus) for comparison.
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Acknowledgement: This subsection of the Thylacine Museum has been referenced (in part) from: ARCHER, M., 1982. A review of Miocene thylacinids (Thylacinidae, Marsupialia), the phylogenetic position of the Thylacinidae and the problem of apriorisms in character analysis. In "Carnivorous Marsupials - Vol. 2" (Ed. M. Archer). Roy. Zool. Soc. N.S.W.: Sydney. pp. 445-76.
References
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